本研究目的主要為探討多種蟹類之胃磨形態與食性差異之間的關連，以及不同的胃磨形態特徵是否能做為辨別蟹種的依據。以台灣西部沿海濕地採集之蟹類為材料，包含13科29屬48種。胃磨主要由位於中央的尾賁門骨及兩側的軛賁門骨組成，是十足目動物用以將攝取的食物研磨成小顆粒的重要工具。本研究將胃磨構造中研磨部位的軛賁門骨、尾賁門骨分別以掃描式電子顯微鏡(Scanning Electron Microscope，SEM )拍照，描述其形態特徵，並比較不同食性之胃磨結構的差異。根據解剖顯微鏡下所觀察的蟹類胃內含物，大致可以歸納出四種類型的螃蟹食性，即濾食型、藻食型、雜食型和肉食型。並且使用主成分分析(Principal Component Analysis ，PCA)，來探討胃磨形態與食性之間的關連，結果發現我們可以用軛賁門骨的形態特徵初步判別蟹種的食性，其中包括軛賁門骨上的梳狀齒數、門齒數及犬齒數。以及用尾賁門骨的尾骨齒板數、軛賁門骨的臼齒數目及前臼齒與臼齒間是否癒合等特定胃磨形態特徵作為分類的依據

The purpose of this study is to investigate the relationship between the morphology of the gastric mill and the difference of feeding habits, as well as whether the characteristics of different gastric mill can be used as the basis for distinguishing crab species. The crabs collected from the western coastal wetlands of Taiwan are composed of 13 species , 29 genera and 48 species. Gastric mill mainly composed by the central of urocardiac ossicle and zygocardiac ossicle on both sides of the, is an important tool of the decapoda to grind the ingested food into small particles. In this study, the urocardiac ossicle and zygocardiac ossicle as the parts of the gastric mill to deal with grinding were taken by scanning electron microscope (SEM), and the morphological characteristics were describe and compared the difference of feeding habit gastric mill morphology . By the contents of the food contained in the stomach contents under the dissecting microscope. The crabs can be roughly classified into four types: the filter-type, the algivore, the omnivorous and the carnivore crabs. The principal component analysis (PCA) was used to investigate the relationship between gastric morphology and feeding habits. The morphological features can be used to determine the feeding habits including the number of comb teeth , incisor teeth and canine teeth on the zygocardiac ossicle. The morphological characteristics such as the number of bone plates of urocardiac ossicle, the number of molars on the zygocardiac ossicle, and the presence or absence of healing between premolar and molars can be used as classification basis.

楊思諒，1986。胃磨結構在蟹類分類研究上的意義 (甲殼綱:歪尾派) 。動物分類學報11: 151-159。
李榮祥，2001。台灣賞蟹情報。大樹文化事業股份有限公司出版，台北。
Allardyce, B. J. & Linton, S.M. (2010) .Functional Morphology of the Gastric Mills of Carnivorous, Omnivorous, and Herbivorous Land Crabs. J Morphol. Jan;271(1):61-72.
Brösing A, Richter S, Scholtz G. 2007. Phylogenetic analysis of the Brachyura (Crustacea, Decapoda) based on characters of the foregut with establishment of a new taxon. J Zool Syst Evol Res 45:20–32.
Brösing , A.(2010). Recent developments on the morphology of the brachyuran foregut ossicles and gastric teeth. Zootaxa 2510(2510):1-44
Brösing A, Türkay M. (2011). Gastric teeth of some thoracotreme crabs and their contribution to the brachyuran phylogeny.
J Morphol 272:1109–1115.
Cannicci, S. et al.(2002). Feeding Habits And Natural Diet of the Intertidal Crab Pachygrapsus marmoratus: Opportunistic Browser or Selective Feeder? Estuar. Coast. Mar. Sci. Volume 54, Issue 6, June 2002, Pages 983-1001
Chisaka H. & Kozawa,Y.(2003). Fine Structure and Mineralization of the Gastric Mill in the Crayfish Procambarus Clarkii During Intermolt Stage. Journal of Crustacean Biology 23:371-379
de Jong L, Jean S, Moreau X (2004) The digestive system of Euphausia krohni (Crustacea, Euphausiacea): functional morphology and phylogenetic relevance. Cah Biol Mar 45:71–82.
Factor, J.R. (1982). Development and metamorphosis of the feeding apparatus of the stone crab, Menippe mercenaria (brachyura, xanthidae). Journal of Morphology 172 (3): 299-312.
Huespe ,A.V., Gómez-Simes, E.& Pastor-de-Ward C.T. (2008). Gastric mill morphology in the genus Cyrtograpsus (Crustacea: Decapoda: Grapsoidea: Varunidae) Volume 88, Issue 2 March , pp. 311-319
Icely, J.D., and Jones , D.A. (1978) Factors affecting the distribution of the genus Uca (Crustacea: Ocypodidae) on an East African shore. Estuarine and Coastal Marine Science 6(3):315–325.
Kobusch W. (1999) The Phylogeny of the Peracarida (Crustacea, Malacostraca). Cuvillier-Verlag, Go¨ ttingen.
Hobbs, K. H. and Hooper , S. L. (2009) High-Resolution Computed Tomography of Lobster(Panulirus interruptus) Stomach. Journal of Morphology 270:1029–1041
Huxley, T. H. (1880) The crayfish: an introduction to the study of zoology. The International Scientific Series, November 1879, 28, 1–371.
Linton,S.M, Allardyce B.J., et al(2009). Food utilisation and digestive ability of aquatic and semi-terrestrial crayfishes, Cherax destructor and Engaeus sericatus (Astacidae, Parastacidae).J Comp Physiol B. May;179(4):493-507.
Maynard, D. M. & Dando, M. R.(1974) The structure of the stomatogastric neuromuscular system in Callinectes sapidus, Homarus americanus, and Panulirus argus (decapoda crustacea). Philos. Trans. R. Soc. Lond. B Biol. Sci. 268, 161-220
Meiss D.E., Norman R.S. (1977). Comparative study of the stomatogastric
system of several decapod crustacea. I. Skeleton. J Morphol 152:21–53.
Milne-Edwards, H. (1834-1837) Histoire naturelle des Crustacés, comprenant l'anatomie, la physiologie et la classification de ces animaux. In: Librairie Encyclopedique de Roret 1, 1–532. Paris.
Mocquard, F. (1883) Recherches anatomiques sur l’estomac des crustacés podophthalmaires. Annales Sciences Naturelles, 16, 1–311.
Parker, T. J. (1876) On the stomach of the fresh-water crayfish. Journal of Anatomy and Physiology, 11, 54–60.
Wa¨ gele JW (1989) Evolution und phylogenetisches System der Isopoda. Zoologica (Stuttgart) 140:1–262.
Sakai K., Tu¨rkay M. and Yang S.L. (2006) Revision of the Helice/
Chasmagnathus complex (Crustacea: Decapoda: Brachyura).
Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft
565, 1–76.
Spears T, Abele LG, Kim W (1992) The monophyly of brachyuran crabs: a phylogenetic study based on 18S rRNA. Syst Biol 41:446–461.
Suthers I.M. & Anderson, A.J. (1981) .Functional morphology of mouthpartsand gastric mill of Ibacus peronii (Leach) (Palinura: Scyllaridae).Aust J Mar Freshw Res 32:931–944
Ter Braak C. J. F., Šmilauer, P.(2002) ANOCO Reference Manual and CanoDraw for Windows User's Guide: Software for Canonical Community Ordination (version 4.5)
Rashid, D.J., Chapman, S.C., et al (2014) From dinosaurs to birds: a tail of evolution. Evodevo. Jul 29
Richter S, Scholtz G (2001) Phylogenetic analysis of the Malacostraca(Crustacea). J Zool Syst Evol Res 39:113–136.